海南岛热带雨林毛蠓亚属（双翅目：蠓科：毛蠓属）2新种及1海南新纪录种

郑诗雅; 李加慧; 林晓丹; 郑诗雅; 李加慧; 林晓丹

doi:10.15886/j.cnki.rdswxb.20250178

[1]	Borkent A, Dominiak P. Catalog of the biting midges of the world (Diptera: Ceratopogonidae) [J]. Zootaxa, 2020, 4787(1): 1−377. https://doi.org/10.11646/zootaxa.4787.1.1 doi: 10.11646/zootaxa.4787.1.1
[2]	Meigen J W. Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten, volume 1 [M]. Aachen: F. W. Forstmann, 1818.(查阅网上资料,请补充引用页码) https://doi.org/10.5962/bhl.title.13731
[3]	Kieffer J J. Beitrag zur Kenntnis der deutschen Ceratopogoninen [J]. Berliner Entomologische Zeitschrift, 1919, 63(4): 201−212. (查阅网上资料, 未找到本条文献信息, 请确认)
[4]	Borkent A, Evenhuis N L, Dominiak P. The world catalog of biting midges (Diptera: Ceratopogonidae): additional updates and errata [J]. Zootaxa, 2025, 5631(2): 245−268. https://doi.org/10.11646/zootaxa.5631.2.2 doi: 10.11646/zootaxa.5631.2.2
[5]	虞以新. 中国蠓科昆虫(昆虫纲, 双翅目)[M]. 北京: 军事医学科学出版社, 2006. (查阅网上资料, 请补充引用页码)
[6]	庄松阳, 彭恒, 袁浩, 等. 中国南沙群岛美济礁蠓种描述(双翅目: 蠓科)[J]. 中国媒介生物学及控制杂志, 2022, 33(5): 732−736. https://doi.org/10.11853/j.issn.1003.8280.2022.05.022 doi: 10.11853/j.issn.1003.8280.2022.05.022
[7]	彭恒, 庄松阳, 袁浩, 等. 中国南沙群岛永暑礁蠓科2新种记述(双翅目: 蠓科)[J]. 中国热带医学, 2023, 23(2): 196−199. https://doi.org/10.13604/j.cnki.46-1064/r.2023.02.18 doi: 10.13604/j.cnki.46-1064/r.2023.02.18
[8]	王崇财, 陈星恒, 谭润谦, 等. 海南蠓科昆虫名录(双翅目: 蠓科)[J]. 中国国境卫生检疫杂志, 2011, 34(1): 24−27. https://doi.org/10.16408/j.1004-9770.2011.01.001 doi: 10.16408/j.1004-9770.2011.01.001
[9]	Grogan W L, Jr, Wieners J A. A new species of the biting midge genus Dasyhelea Kieffer (Diptera: Ceratopogonidae) from the Bahamas [J]. Proceedings of the Entomological Society of Washington, 2006, 108(2): 467−473.
[10]	Dominiak P. Biting midges of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae) in Poland [J]. Polish Journal of Entomology, 2012, 81(3): 211−304. https://doi.org/10.2478/v10200-012-0009-8 doi: 10.2478/v10200-012-0009-8
[11]	Waugh W T, Wirth W W. A revision of the genus Dasyhelea Kieffer of the eastern United States north of Florida (Diptera: Ceratopogonidae) [J]. Annals of the Entomological Society of America, 1976, 69(2): 219−247. https://doi.org/10.1093/aesa/69.2.219 doi: 10.1093/aesa/69.2.219
[12]	Graves R C, Graves A C F. Diptera associated with shelf fungi and certain other micro-habitats in the highlands area of western North Carolina [J]. Entomological News, 1985, 96(2): 87−92.
[13]	Vattier G. Dasyhelea adami sp. n. (Diptera, Ceratopogonidae) morphologie et biologie [J]. Bulletin de la Société de Pathologie Exotique, 1964, 57(5): 1159−1177.
[14]	Johannsen O A. Ceratopogoninae from the Malayan subregion of the Dutch East Indies [J]. Archiv für Hydrobiologie, 1931, S9: 403−448.
[15]	Mayer K. Die Metamorphose der Ceratopogonidae (Dipt. ). Ein Beitrag zur Morphologie, Systematik, Ökologie und Biologie der Jugendstadien dieser Dipterenfamilie [J]. Archiv für Naturgeschichte, 1934, 3: 205−288.
[16]	Mayer K. Ceratopogoniden-metamorphosen (C. intermediae und C. vermiformes) der deutschen limnologischen Sunda-expedition [J]. Archiv für Hydrobiologie, 1934, S13: 166−202.
[17]	Zilahi-Sebess G. Über heleiden des Balaton-gebiets [J]. Arbeiten des Ungarischen Biologischen Forschungs-Institutes, Tihany, 1936, 8: 196−206.
[18]	Przhiboro A A. New data on Ceratopogonidae from lake shore habitats in NW Russia (Diptera) [J]. Zoosystematica Rossica, 2005, 14(1): 165−167. https://doi.org/10.31610/zsr/2005.14.1.165 doi: 10.31610/zsr/2005.14.1.165
[19]	Dominiak P, Szadziewski R. Biting midges of the genus Dasyhelea Kieffer, 1911 (Diptera: Ceratopogonidae) collected from Polish peat-bogs [J]. Dipteron, 2006, 22: 4−7.
[20]	Lee K M, Chan K L. The biology of Dasyhelea ampullariae in monkey cups at Kent Ridge (Diptera: Ceratopogonidae) [J]. Journal of the Singapore National Academy of Science, 1985, 14: 6−14.
[21]	Bogarín D, Fernández M, Borkent A, et al. Pollination of Trichosalpinx (Orchidaceae: Pleurothallidinae) by biting midges (Diptera: Ceratopogonidae) [J]. Botanical Journal of the Linnean Society, 2018, 186(4): 510−543. https://doi.org/10.1093/botlinnean/box087 doi: 10.1093/botlinnean/box087
[22]	Kutty S N, Wong W H, Meusemann K, et al. A phylogenomic analysis of Culicomorpha (Diptera) resolves the relationships among the eight constituent families [J]. Systematic Entomology, 2018, 43(3): 434−446. https://doi.org/10.1111/syen.12285 doi: 10.1111/syen.12285
[23]	Lee K M, Wirth W W, Chan K L. A new species of Dasyhelea midge reared from drains in Singapore (Diptera: Ceratopogonidae) [J]. Proceedings of the Entomological Society of Washington, 1989, 91(3): 452−457.
[24]	Vandromme M, Van de Sande E, Pinceel T, et al. Resolving the identity and breeding habitats of cryptic dipteran cacao flower visitors in a neotropical cacao agroforestry system [J]. Basic and Applied Ecology, 2023, 68: 35−45. https://doi.org/10.1016/j.baae.2023.03.002 doi: 10.1016/j.baae.2023.03.002
[25]	Ríos-Moyano D K, Rodríguez-Cruz F A, Hormaza-Martínez P A, et al. Characterization of pollinators associated with cocoa cultivation and their relationship with natural effective pollination [J]. Diversity, 2025, 17(3): 189. https://doi.org/10.3390/d17030189 doi: 10.3390/d17030189
[26]	Winder J A. Ecology of insects in the cocoa environment [J]. Annual Review of Entomology, 1978, 23(1): 123−126. (查阅网上资料, 未找到本条文献信息, 请确认)
[27]	吴小香, 杨浩, 孟祥瑞, 等. 阿蠓属一新种及一中国新记录种(昆虫纲: 双翅目: 蠓科)[J]. 环境昆虫学报, 2022, 44(4): 1043−1052. https://doi.org/10.3969/j.issn.1674-0858.2022.04.30 doi: 10.3969/j.issn.1674-0858.2022.04.30
[28]	孟祥瑞, 李加慧, 林晓丹. 海南岛铗蠓属尤蠓亚属(双翅目: 蠓科)一新种及一海南新纪录种[J]. 热带生物学报, 2025, 16(4): 553−560. https://doi.org/10.15886/j.cnki.rdswxb.20240071 doi: 10.15886/j.cnki.rdswxb.20240071
[29]	Wirth W W, Hubert A A. The Culicoides of Southeast Asia (Diptera: Ceratopogonidae) [M]. Gainesville: American Entomological Institute, 1989: 40−453.
[30]	虞以新. 中国蠓科昆虫: 昆虫纲, 双翅目(第1、2卷)[M]. 北京: 军事医学科学出版社, 2006: 816−1322. (查阅网上资料, 请确认与第5条文献是否重复)
[31]	Borkent A. Ceratopogonidae[M]//Kirk-Spriggs A H, Sinclair B J. Manual of Afrotropical Diptera. Volume 2. Nematocerous Diptera and lower Brachycera: Suricata 5. Pretoria: South African National Biodiversity Institute, 2017: 733−812.
[32]	Tamura K, Stecher G, Kumar S. MEGA11: molecular evolutionary genetics analysis version 11 [J]. Molecular Biology and Evolution, 2021, 38(7): 3022−3027. https://doi.org/10.1093/molbev/msab120 doi: 10.1093/molbev/msab120
[33]	Kimura M. A simple method for estimating evolutionary rates of base substitutions through comparative studies of nucleotide sequences [J]. Journal of Molecular Evolution, 1980, 16(2): 111−120. https://doi.org/10.1007/BF01731581 doi: 10.1007/BF01731581
[34]	虞以新. 我国香港特别行政区蠓类二新种及新记录种的发现(双翅目: 蠓科)[J]. 寄生虫与医学昆虫学报, 2001, 8(3): 159−163.
[35]	Tokunaga M. Biting midges of the genus Dasyhelea from Japan [J]. Publications of the Entomological Laboratory, College of Agriculture, University of Kyoto, 1962, 12: 1−25. (查阅网上资料, 未找到本条文献信息, 请确认)
[36]	Wirth W W. A review of the species of the genus Dasyhelea from the Greater Antilles [J]. Transactions of the American Entomological Society, 1971, 97(2): 209−235. (查阅网上资料, 未找到本条文献信息, 请确认)