[1] Yruela I. Copper in plants: acquisition, transport and interactions [J]. Functional Plant Biology, 2009, 36(5): 409−430. https://doi.org/10.1071/fp08288 doi:  10.1071/fp08288
[2] Ravet K, Pilon M. Copper and iron homeostasis in plants: the challenges of oxidative stress [J]. Antioxidants & Redox Signaling, 2013, 19(9): 919−932. https://doi.org/10.1089/ars.2012.5084 doi:  10.1089/ars.2012.5084
[3] Rai S, Singh P K, Mankotia S, et al. Iron homeostasis in plants and its crosstalk with copper, zinc, and manganese [J]. Plant Stress, 2021, 1: 100008. https://doi.org/10.1016/j.stress.2021.100008 doi:  10.1016/j.stress.2021.100008
[4] Marschner P. Marschner’s mineral nutrition of higher plants [M]. 3rd ed. London: Elsevier/Academic Press, 2012: 651. doi: 10.1016/C2009-0-63043-9
[5] Sancenón V, Puig S, Mateu-Andrés I, et al. The Arabidopsis copper transporter COPT1 functions in root elongation and pollen development [J]. Journal of Biological Chemistry, 2004, 279(15): 15348−15355. https://doi.org/10.1074/jbc.M313321200 doi:  10.1074/jbc.M313321200
[6] Lequeux H, Hermans C, Lutts S, et al. Response to copper excess in Arabidopsis thaliana: impact on the root system architecture, hormone distribution, lignin accumulation and mineral profile [J]. Plant Physiology and Biochemistry, 2010, 48(8): 673−682. https://doi.org/10.1016/j.plaphy.2010.05.005 doi:  10.1016/j.plaphy.2010.05.005
[7] Adrees M, Ali S, Rizwan M, et al. The effect of excess copper on growth and physiology of important food crops: a review [J]. Environmental Science and Pollution Research, 2015, 22(11): 8148−8162. https://doi.org/10.1007/s11356-015-4496-5 doi:  10.1007/s11356-015-4496-5
[8] Cobbett C, Goldsbrough P. Phytochelatins and metallothioneins: roles in heavy metal detoxification and homeostasis [J]. Annual Review of Plant Biology, 2002, 53: 159−182. https://doi.org/10.1146/annurev.arplant.53.100301.135154 doi:  10.1146/annurev.arplant.53.100301.135154
[9] Peñarrubia L, Andrés-Colás N, Moreno J, et al. Regulation of copper transport in Arabidopsis thaliana: a biochemical oscillator? [J]. JBIC Journal of Biological Inorganic Chemistry, 2010, 15(1): 29−36. https://doi.org/10.1007/s00775-009-0591-8 doi:  10.1007/s00775-009-0591-8
[10] Deng F L, Yamaji N, Xia J X, et al. A member of the heavy metal P-type ATPase OsHMA5 is involved in xylem loading of copper in rice [J]. Plant Physiology, 2013, 163(3): 1353−1362. https://doi.org/10.1104/pp.113.226225 doi:  10.1104/pp.113.226225
[11] Carrió-Seguí À, Romero P, Curie C, et al. Copper transporter COPT5 participates in the crosstalk between vacuolar copper and iron pools mobilisation [J]. Scientific Reports, 2019, 9(1): 4648. https://doi.org/10.1038/s41598-018-38005-4 doi:  10.1038/s41598-018-38005-4
[12] Thounaojam T C, Panda P, Mazumdar P, et al. Excess copper induced oxidative stress and response of antioxidants in rice [J]. Plant Physiology and Biochemistry, 2012, 53: 33−39. https://doi.org/10.1016/j.plaphy.2012.01.006 doi:  10.1016/j.plaphy.2012.01.006
[13] 谢晓霄, 徐劼, 方学雷, 等. 铜胁迫下吊兰的生理响应及对铜的吸收特性[J]. 广州化工, 2023, 51(10): 41−43. https://doi.org/10.3969/j.issn.1001-9677.2023.10.014 doi:  10.3969/j.issn.1001-9677.2023.10.014
[14]

Sposito G. The chemistry of soils [M]. 2nd ed. Oxford: Oxford University Press, 2008: 329.
[15] 李明谦, 张凤良, 李小琴, 等. 橡胶树无性系苗期生长和光合生理特性的比较[J]. 西南林业大学学报(自然科学), 2020, 40(2): 18−22. https://doi.org/10.11929/j.swfu.201905032 doi:  10.11929/j.swfu.201905032
[16] 贝美容, 蔡隽, 戚春林, 等. 2种新型种植材料(苗木)橡胶幼树的施肥效应[J]. 热带作物学报, 2019, 40(11): 2105−2111. https://doi.org/10.3969/j.issn.1000-2561.2019.11.002 doi:  10.3969/j.issn.1000-2561.2019.11.002
[17] 张婷婷, 刘子凡, 安锋, 等. 铝胁迫造成橡胶苗死亡的机制研究[J]. 热带作物学报, 2020, 41(12): 2439−2445. https://doi.org/10.3969/j.issn.1000-2561.2020.12.010 doi:  10.3969/j.issn.1000-2561.2020.12.010
[18]

Bolan N S, Adriano D C, Curtin D. Soil acidification and liming interactions with nutrientand heavy metal transformationand bioavailability [J]. Advances in Agronomy, 2003, 78: 215−272. https://doi.org/10.1016/s0065-2113(02)78006-1 doi:  10.1016/s0065-2113(02)78006-1
[19]

Liu C A, Liang M Y, Nie Y, et al. The conversion of tropical forests to rubber plantations accelerates soil acidification and changes the distribution of soil metal ions in topsoil layers [J]. Science of the Total Environment, 2019, 696: 134082. https://doi.org/10.1016/j.scitotenv.2019.134082 doi:  10.1016/j.scitotenv.2019.134082
[20] 覃怀德, 高乐, 吴炳孙, 等. 橡胶树割面涂施硫酸铜对胶乳生理特性影响的初步分析[J]. 热带农业科学, 2013, 33(6): 1−4. https://doi.org/10.3969/j.issn.1009-2196.2013.06.001 doi:  10.3969/j.issn.1009-2196.2013.06.001
[21] 肖再云, 校现周. 橡胶树胶乳铜离子含量与乙烯刺激强度的关系初探[J]. 热带农业科技, 2010, 33(3): 10−12. https://doi.org/10.3969/j.issn.1672-450X.2010.03.003 doi:  10.3969/j.issn.1672-450X.2010.03.003
[22]

Hoagland D R, Arnon D I. The water-culture method for growing plants without soil [M]. Berkeley: The College of Agriculture, University of California, 1950: 32.
[23] 韩志平, 张海霞, 赵智灵, 等. 铜胁迫对小白菜幼苗生长的影响[J]. 长江蔬菜, 2015(12): 28−32. https://doi.org/10.3865/j.issn.1001-3547.2015.12.010 doi:  10.3865/j.issn.1001-3547.2015.12.010
[24] 陆奇杰, 巢建国, 谷巍, 等. 铜胁迫对茅苍术光合特性及生理指标的影响[J]. 中药材, 2018, 41(7): 1534−1538. https://doi.org/10.13863/j.issn1001-4454.2018.07.004 doi:  10.13863/j.issn1001-4454.2018.07.004
[25] 张筱, 张海芹, 高宇琪, 等. 不同地被覆盖物对干旱胁迫下银杏生长及抗旱性的影响[J]. 贵州农业科学, 2025, 53(12): 135−144. https://doi.org/10.3969/j.issn.1001-3601.2025.12.015 doi:  10.3969/j.issn.1001-3601.2025.12.015
[26] 李辉容, 唐天君, 杨文亮. 火焰原子吸收光谱法测定铜锤草中锰铁铜锌[J]. 食品与发酵科技, 2013, 49(3): 72−73. https://doi.org/10.3969/j.issn.1674-506X.2013.03-020 doi:  10.3969/j.issn.1674-506X.2013.03-020
[27] 郭英. 火焰原子吸收分光光度法测定地表水中的铜、锌、铅、镉[J]. 环境科学导刊, 2017, 36(3): 106−108. https://doi.org/10.13623/j.cnki.hkdk.2017.03.022 doi:  10.13623/j.cnki.hkdk.2017.03.022
[28] 张艳, 李勋, 彭彬, 等. 雅砻江上游甘孜段历史遗留矿山乡土植物重金属富集能力比较[J]. 应用与环境生物学报, 2023, 29(6): 1373−1380. https://doi.org/10.19675/j.cnki.1006-687x.2022.10006 doi:  10.19675/j.cnki.1006-687x.2022.10006
[29] 孟桂元, 蒋文杰, 韩杰铖, 等. 锑胁迫对苎麻生长及锑富集转运的影响[J]. 生态学杂志, 2026, 45(5): 1577−1585. https://doi.org/10.13292/j.1000-4890.202605.015 doi:  10.13292/j.1000-4890.202605.015
[30]

Yamasaki H, Hayashi M, Fukazawa M, et al. SQUAMOSA promoter binding Protein-Like7 is a central regulator for copper homeostasis in Arabidopsis [J]. The Plant Cell, 2009, 21(1): 347−361. https://doi.org/10.1105/tpc.108.060137 doi:  10.1105/tpc.108.060137
[31]

Hayat S, Hayat Q, Alyemeni M N, et al. Role of proline under changing environments: a review [J]. Plant Signaling & Behavior, 2012, 7(11): 1456−1466. https://doi.org/10.4161/psb.21949 doi:  10.4161/psb.21949
[32]

Szabados L, Savouré A. Proline: a multifunctional amino acid [J]. Trends in Plant Science, 2010, 15(2): 89−97. https://doi.org/10.1016/j.tplants.2009.11.009 doi:  10.1016/j.tplants.2009.11.009
[33]

Dar M I, Naikoo M I, Rehman F, et al. Proline accumulation in plants: roles in stress tolerance and plant development[M]//Iqbal N, Nazar R, Khan N A. Osmolytes and plants acclimation to changing environment: emerging omics technologies. New Delhi: Springer, 2016: 155−166. https://doi.org/10.1007/978-81-322-2616-1_9
[34]

Sancenón V, Puig S, Mira H, et al. Identification of a copper transporter family in Arabidopsis thaliana [J]. Plant Molecular Biology, 2003, 51(4): 577−587. https://doi.org/10.1023/A:1022345507112 doi:  10.1023/A:1022345507112