[1] DESNEUX N, WAJNBERG E, WYCKHUYS K A G, et al. Biological invasion of European tomato crops by Tuta absoluta: ecology, geographic expansion and prospects for biological control [J]. Journal of Pest Science, 2010, 83(3): 197 − 215. doi:  10.1007/s10340-010-0321-6
[2] 张桂芬, 刘万学, 万方浩, 等. 世界毁灭性检疫害虫番茄潜叶蛾的生物生态学及危害与控制[J]. 生物安全学报, 2018, 27(3): 155 − 163. doi:  10.3969/j.issn.2095-1787.2018.03.001
[3]

VISSER D, UYS V M, NIEUWENHUIS R J, et al. First records of the tomato leaf miner Tuta absoluta (Meyrick, 1917) (Lepidoptera: Gelechiidae) in South Africa [J]. BioInvasions Records, 2017, 6(4): 301 − 305. doi:  10.3391/bir.2017.6.4.01
[4]

SANTANA P A, KUMAR L, SILVA R S, et al. Global geographic distribution of Tuta absoluta as affected by climate change [J]. Journal of Pest Science, 2019, 92(4): 1373 − 1385. doi:  10.1007/s10340-018-1057-y
[5]

FAO (Food and Agriculture Organization). Online statistical database: food and agriculture data. FAOSTAT [EB/OL]. (2012) [2022-09-23]. http://www.fao.org/faostat.
[6]

FAO (Food and Agriculture Organization). Online statistical database: food and agriculture data. FAOSTAT [EB/OL]. (2017) [2022-09-23]. http://www.fao.org/faostat.
[7]

XIAN X Q, HAN P, WANG S, et al. The potential invasion risk and preventive measures against the tomato leafminer Tuta absoluta in China [J]. Entomologia Generalis, 2017, 36(4): 319 − 333. doi:  10.1127/entomologia/2017/0504
[8] 冼晓青, 张桂芬, 刘万学, 等. 世界性害虫番茄潜麦蛾入侵我国的风险分析[J]. 植物保护学报, 2019, 46(1): 49 − 55. doi:  10.13802/j.cnki.zwbhxb.2019.2019906
[9] 张桂芬, 马德英, 刘万学, 等. 中国新发现外来入侵害虫—南美番茄潜叶蛾(鳞翅目: 麦蛾科)[J]. 生物安全学报, 2019, 28(3): 200 − 203. doi:  10.3969/j.issn.2095-1787.2019.03.007
[10] 张桂芬, 冼晓青, 张毅波, 等. 警惕南美番茄潜叶蛾Tuta absoluta (Meyrick)在中国扩散[J]. 植物保护, 2020, 46(2): 281 − 286.
[11]

CABI. Invasive species compendium Phthorimaea absoluta (tomato leafminer) datasheet [EB/OL]. (2020-11-30) [2022-09-23]. https://www.cabi.org/isc/datasheet/49260#: ～:text= Phthorimaea absoluta.
[12]

BIONDI A, GUEDES R N C, WAN F H, et al. Ecology, worldwide spread, and management of the invasive South American tomato pinworm, Tuta absoluta: past, present, and future [J]. Annual Review of Entomology, 2018, 63: 239 − 258. doi:  10.1146/annurev-ento-031616-034933
[13] 张桂芬, 张毅波, 刘万学, 等. 诱捕器颜色和悬挂高度对番茄潜叶蛾诱捕效果的影响[J]. 中国农业科学, 2021, 54(11): 2343 − 2354. doi:  10.3864/j.issn.0578-1752.2021.11.008
[14]

SALAZAR E R, ARAYA J E. Detection of insecticide resistance in the tomato moth [J]. Simiente, 1997, 67: 8 − 22.
[15]

SIQUEIRA H A A, GUEDES R N C, PICANçO M C. Insecticide resistance in populations of Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Agriculture and Forest Entomology, 2000, 2(2): 147 − 153. doi:  10.1046/j.1461-9563.2000.00062.x
[16]

LIETTI M M M, BOTTO E, ALZOGARAY R A. Insecticide resistance in Argentine populations of Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. Neotropical Entomology, 2005, 34: 113 − 119. doi:  10.1590/S1519-566X2005000100016
[17]

GUEDES R N C, PICANçO M C. The tomato borer Tuta absoluta in South America: pest status, management and insecticide resistance [J]. Bulletin OEPP/EPPO Bulletin, 2012, 42(2): 211 − 216. doi:  10.1111/epp.2557
[18]

SALAZAR E R, ARAYA J E. Response of the tomato moth, Tuta absoluta (Meyrick), to insecticides in Arica, Chile [J]. Agricultura Tecnica, 2001, 61(4): 429 − 435.
[19]

SIQUEIRA H A A, GUEDES R N C, FRAGOSO D B, et al. Abamectin resistance and synergism in Brazilian populations of Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. International Journal of Pest Management, 2001, 47(4): 247 − 251. doi:  10.1080/09670870110044634
[20]

LUNA M G, SáNCHEZ N E, PEREYRA P C, et al. Biological Control of Tuta absoluta in Argentina and Italy: evaluation of indigenous insects as natural enemies [J]. Bulletin OEPP/EPPO Bulletin, 2012, 42(2): 260 − 267. doi:  10.1111/epp.2564
[21]

JOSHI A, THAPA R B, KALAUNI D. Integrated management of South American tomato leaf miner [Tuta absoluta (Meyrick)]: a review [J]. Journal of the Plant Protection Society, 2018, 5: 70 − 86. doi:  10.3126/jpps.v5i0.37759
[22]

CABELLO T, GALLEGO J R, VILA E, et al. Biological Control of the South American tomato pinworm, Tuta absoluta (Lep. : Gelechiidae), with releases of Trichogramma achaeae (Hym. : Trichogrammatidae) in tomato greenhouses of Spain [J]. IOBC/WPRS Bulletin, 2009, 49: 225 − 230.
[23]

ZHANG Y B, TIAN X C, WANG H, et al. Host selection behavior of the host-feeding parasitoid Necremnus tutae on Tuta absoluta [J]. Entomologia Generalis, 2021, 42(3): 445 − 456.
[24]

ZAPPALà L, BIONDI A, ALMA A, et al. Natural enemies of the South American moth, Tuta absoluta, in Europe, North Africa and Middle East, and their potential use in pest control strategies [J]. Journal of Pest Science, 2013, 86(4): 635 − 647. doi:  10.1007/s10340-013-0531-9
[25]

VAN LENTEREN J C, BUENO V H P, BURGIO G, et al. Pest kill rate as aggregate evaluation criterion to rank biological control agents: a case study with Neotropical predators of Tuta absoluta on tomato [J]. Bulletin of Entomological Research, 2019, 109(6): 812 − 820. doi:  10.1017/S0007485319000130
[26]

CABELLO T, GALLEGO J R, FERNáNDEZ F J, et al. Biological Control strategies for the South American tomato moth (Lepidoptera: Gelechiidae) in greenhouse Tomatoes [J]. Journal of Economic Entomology, 2012, 105(6): 2085 − 2096. doi:  10.1603/EC12221
[27]

BIONDI A, CHAILLCUX A, LAMBION J, et al. Indigenous natural enemies attacking Tuta absoluta (Lepidoptera: Gelechiidae) in Southern France [J]. Egyptian Journal of Biological Pest Control, 2013, 23(1): 117 − 121.
[28]

OLIVEIRA L, DURãO A C, FONTES J, et al. Potential of Trichogramma achaeae (Hymenoptera: Trichogrammatidae) in Biological Control of Tuta absoluta (Lepidoptera: Gelechiidae) in Azorean Greenhouse Tomato Crops [J]. Journal of Economic Entomology, 2017, 110(5): 2010 − 2015. doi:  10.1093/jee/tox197
[29]

MOHAMADI P, RAZMJOU J, NASERI B, et al. Humic fertilizer and vermicompost applied to the soil can positively affect population growth parameters of Trichogramma brassicae (Hymenoptera: Trichogrammatidae) on eggs of Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Neotropical Entomology, 2017, 46(6): 678 − 684. doi:  10.1007/s13744-017-0536-9
[30]

SARHAN A A, OSMAN M A M, MANDOUR N S, et al. Parasitization capability of four trichogrammatid species against the tomato leaf miner, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) under different releasing regimes [J]. Egyptian Journal of Biological Pest Control, 2015, 25(1): 107 − 112.
[31]

ABBES K, HARBI A, CHERMITI B. The tomato leafminer Tuta absoluta (Meyrick) in Tunisia: current status and management strategies [J]. Bulletin OEPP/EPPO Bulletin, 2012, 42(2): 226 − 233. doi:  10.1111/epp.2559
[32]

EL-ARNAOUTY S A, PIZZOL J, GALAL H H, et al. Assessment of two Trichogramma species for the control of Tuta absoluta in North African tomato greenhouses [J]. African Entomology, 2014, 22(4): 801 − 809. doi:  10.4001/003.022.0410
[33]

ALSAEDI G, ASHOURI A, TALAEI-HASSANLOUI R. Assessment of two Trichogramma species with Bacillus thuringiensis var. krustaki for the control of the tomato leafminer Tuta absoluta Meyrick (Lepidoptera: Gelechiidae) in Iran [J]. Open Journal of Ecology, 2017, 7(2): 112 − 124. doi:  10.4236/oje.2017.72009
[34]

CHAILLEUX A, BEAREZ P, PIZZOL J, et al. Potential for combined use of parasitoids and generalist predators for biological control of the key invasive tomato pest Tuta absoluta [J]. Journal of Pest Science, 2013, 86(3): 533 − 541. doi:  10.1007/s10340-013-0498-6
[35]

CALVO F J, SORIANO J D, STANSLY P A, et al. 2016. Can the parasitoid Necremnus tutae (Hymenoptera: Eulophidae) improve existing biological control of the tomato leafminer Tuta absoluta (Lepidoptera: Gelechiidae) ? [J]. Bulletin of Entomological Research, 2016, 1(4): 1 − 10.
[36]

FERRACINI C, INGEGNO B L, NAVONE P, et al. Adaptation of indigenous larval parasitoids to Tuta absoluta (Lepidoptera: Gelechiidae) in Italy [J]. Journal of Economic Entomology, 2012, 105(4): 1311 − 1319. doi:  10.1603/EC11394
[37]

ZAPPALà L, BERNARDO U, BIONDI A, et al. Recruitment of native parasitoids by the exotic pest Tuta absoluta in Southern Italy [J]. Bulletin of Insectology, 2012, 65(1): 51 − 61.
[38]

BAYRAM Y, GüLER Y, FURSOV V, et al. First record of Necremnus cosmopterix Ribeset Bernardo, 2015 (Hymenoptera: Eulophidae), as a larval parasitoid of tomato leaf miner, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) in Turkey [J]. Egyptian Journal of Biological Pest Control, 2016, 26(4): 847 − 848.
[39]

URBANEJA A, GONZALEZ-CABRERA J, ARNó J, et al. Prospects for the biological control of Tuta absoluta in tomatoes of the Mediterranean basin [J]. Pest Management Science, 2012, 68(9): 1215 − 1222. doi:  10.1002/ps.3344
[40]

CHAILLEUX A, DROUI A, BEAREZ P, et al. Survival of a specialist natural enemy experiencing resource competition with an omnivorous predator when sharing the invasive prey Tuta absoluta [J]. Ecology and Evolution, 2017, 7(20): 8329 − 8337. doi:  10.1002/ece3.3396
[41]

GABARRA R, ARNó J, LARA L, et al. Native parasitoids associated with Tuta absoluta in the tomato production areas of the spanish Mediterranean Coast [J]. BioControl, 2014, 59(1): 45 − 54. doi:  10.1007/s10526-013-9545-8
[42]

YARAHMADI F, SALEHI Z, LOTFALIZADEH H. Two species of the genus Elachertus Spinola (Hym. : Eulophidae) new larval ectoparasitoids of Tuta absoluta (Meyreck) (Lep. : Gelechidae) [J]. Journal of Crop Protection, 2016, 5(3): 413 − 418. doi:  10.18869/modares.jcp.5.3.413
[43]

MELO M, CAMPOS A D. Occurrence of natural enemies on tomato moth Tuta absoluta (Meirick, 1917) (Lepidoptera, Gelechiidae) in Pelotas, Rio Grande do Sul [J]. Agropecuaria Clima Temperado, 2000, 3(2): 269 − 274.
[44]

LUNA M G, SáNCHEZ N E, PEREYRA P C. Parasitism of Tuta absoluta (Lepidoptera, Gelechiidae) by Pseudapanteles dignus (Hymenoptera, Braconidae) under laboratory conditions [J]. Environmental Entomology, 2007, 36(4): 887 − 893. doi:  10.1093/ee/36.4.887
[45]

BIONDI A, DESNEUX N, AMIENS-DESNEUX E, et al. Biology and developmental strategies of the Palaearctic parasitoid Bracon nigricans (Hymenoptera: Braconidae) on the neotropical moth Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Journal of Economic Entomology, 2013, 106(4): 1638 − 1647. doi:  10.1603/EC12518
[46]

IDRISS G E A, MOHAMED S A, KHAMIS F, et al. Biology and performance of two indigenous larval parasitoids on Tuta absoluta (Lepidoptera: Gelechiidae) in Sudan [J]. Biocontrol Science and Technology, 2018, 28(6): 614 − 628. doi:  10.1080/09583157.2018.1477117
[47]

BALLAL C R, GUPTA A, MOHAN M, et al. The new invasive pest Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) in India and its natural enemies along with evaluation of Trichogrammatids for its Biological Control [J]. Current Science, 2016, 110(11): 2155 − 2159. doi:  10.18520/cs/v110/i11/2155-2159
[48]

PARRA J R P, ZUCCHI R A. Trichogramma in Brazil: feasibility of use after twenty years of research [J]. Neotropical Entomology, 2004, 33(3): 271 − 281. doi:  10.1590/S1519-566X2004000300001
[49]

CHAILLEUX A, DESNEUX N, SEGURET J, et al. Assessing European egg parasitoids as a mean of controlling the invasive South American tomato pinworm Tuta absoluta [J]. PLoS ONE, 2012, 7(10): e48068. doi:  10.1371/journal.pone.0048068
[50]

CHAILLEUX A, BIONDI A, HAN P, et al. Suitability of the pest-plant system Tuta absoluta (Lepidoptera: Gelechiidae) - tomato for Trichogramma (Hymenoptera: Trichogrammatidae) parasitoids and insights for Biological Control [J]. Journal of Economic Entomology, 2013, 106(6): 2310 − 2321. doi:  10.1603/EC13092
[51]

GODA N F, EL-HENEIDY A H, DJELOUAH K, et al. Integrated pest management of the tomato leaf miner, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) in tomato fields in Egypt [J]. Egyptian Journal of Biological Pest Control, 2015, 25(3): 655 − 661.
[52]

CABELLO T, GáMEZ M, VARGA Z, et al. Selection of Trichogramma spp. (Hym. : Trichogrammatidae) for the biological control of Tuta absoluta (Lep. : Gelechiidae) in greenhouses by an entomo-ecological simulation model [J]. IOBC/WPRS Bulletin, 2012, 80: 171 − 176.
[53]

CALVO F J, SORIANO J D, BOLCKMANS K, et al. Host instar suitability and life-history parameters under different temperature regimes of Necremnus artynes on Tuta absoluta [J]. Biocontrol Science and Technology, 2013, 23(7): 803 − 815. doi:  10.1080/09583157.2013.795930
[54]

BODINO N, FERRACINI C, TAVELLA L. Is host selection influenced by natal and adult experience in the parasitoid Necremnus tutae (Hymenoptera: Eulophidae)? [J]. Animal Behaviour, 2016, 112: 221 − 228. doi:  10.1016/j.anbehav.2015.12.011
[55]

BODINO N, FERRACINI C, TAVELLA L. Functional response and age-specific foraging behavior of Necremnus tutae and N. cosmopterix, native natural enemies of the invasive pest Tuta absoluta in Mediterranean area [J]. Journal of Pest Science, 2019, 92: 1467 − 1478. doi:  10.1007/s10340-018-1025-6
[56]

CRISOL-MARTíNEZ E, VAN DER BLOM J. Necremnus tutae (Hymenoptera, Eulophidae) is widespread and efficiently controls Tuta absoluta in tomato greenhouses in SE Spain [J]. IOBC/WPRS Bulletin, 2019, 147: 22 − 29.
[57]

GULERIA P, SHARMA P L, VERMA S C, et al. Life history traits and host-killing rate of Neochrysocharis formosa on Tuta absoluta [J]. BioControl, 2020, 65(9): 401 − 411.
[58]

LUNA M G, WADA V I, LA SALLE J, et al. Neochrysocharis formosa (Westwood) (Hymenoptera: Eulophidae), a newly recorded parasitoid of the tomato moth, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae), in Argentina [J]. Neotropical Entomology, 2011, 40(3): 412 − 414. doi:  10.1590/S1519-566X2011000300019
[59]

GEBIOLA M, BERNARDO U, RIBES A, et al. An integrative study of Necremnus Thomson (Hymenoptera: Eulophidae) associated with invasive pests in Europe and North America: taxonomic and ecological implications [J]. Zoological Journal of the Linnean Society, 2015, 173(2): 352 − 423. doi:  10.1111/zoj.12210
[60]

ARNò J, OVEJA M F, GABARRA R. Selection of flowering plants to enhance the biological control of Tuta absoluta using parasitoids [J]. Biological Control, 2018, 122: 41 − 50. doi:  10.1016/j.biocontrol.2018.03.016
[61]

SáNCHEZ N E, PEREYRA P C, LUNA M G. Spatial patterns of parasitism of the solitary parasitoid Pseudapanteles dignus (Hymenoptera: Braconidae) on Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Environmental Entomology, 2009, 38(2): 365 − 374. doi:  10.1603/022.038.0208
[62]

NIEVES E L, PEREYRA P C, LUNA M G, et al. Laboratory population parameters and field impact of the larval endoparasitoid Pseudapanteles dignus (Hymenoptera: Braconidae) on its host Tuta absoluta (Lepidoptera: Gelechiidae) in tomato crops in Argentina [J]. Journal of Economic Entomology, 2015, 108(4): 1553 − 1559. doi:  10.1093/jee/tov115
[63]

URBANEJA A, MONTóN H, MOLLá ó. Suitability of the tomato borer Tuta absoluta as prey for Macrolophus pygmaeus and Nesidiocoris tenuis [J]. Journal of Applied Entomology, 2009, 133(4): 292 − 296. doi:  10.1111/j.1439-0418.2008.01319.x
[64]

JAWORSKI C C, BOMPARD A, GENIES L, et al. Preference and prey switching in a generalist predator attacking local and invasive alien pests [J]. PLoS ONE, 2013, 8(12): e82231. doi:  10.1371/journal.pone.0082231
[65]

LINS J C, LOON J J A, VAN BUENO V H P, et al. Response of the zoophytophagous predators Macrolophus pygmaeus and Nesidiocoris tenuis to volatiles of uninfested plants and to plants infested by prey or conspecifics [J]. BioControl, 2014, 59(6): 707 − 718. doi:  10.1007/s10526-014-9602-y
[66]

VAN LENTEREN J C, VAN HEMERIK L, LINS J C, et al. Functional responses of three Neotropical mirid predators to eggs of Tuta absoluta on tomato [J]. Insects, 2016, 7(3): 34. doi:  10.3390/insects7030034
[67]

SILVA D B, BUENO V H P, MONTES F C, et al. Population growth of three mirid predatory bugs feeding on eggs and larvae of Tuta absoluta on tomato [J]. BioControl, 2016, 61(5): 545 − 553. doi:  10.1007/s10526-016-9736-1
[68]

BUENO V H P, MONTES F C, PEREIRA A M C, et al. Can recently found Brazilian hemipteran predatory bugs control Tuta absoluta? [J]. IOBC/WPRS Bulletin, 2012, 80: 63 − 67.
[69]

BAYRAM Y, BEKTAŞ ö, BüYüK M, et al. A survey of tomato leafminer [(Tuta absoluta Meyrick) (Lepidoptera: Gelechiidae)] and its natural enemies in the South-east Anatolia Region [J]. Turkiye Biyolojik Mucadele Dergisi, 2014, 5(2): 99 − 110.
[70]

QUEIROZ O S, RAMOS R S, GONTIJO L M, et al. Functional response of three species of predatory pirate bugs attacking eggs of Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Environmental Entomology, 2015, 44(2): 246 − 251. doi:  10.1093/ee/nvu026
[71]

MIRANDA M M M, PICANçO M C, ZANUNCIO J C, et al. Impact of integrated pest management on the population of leafminers, fruit borers, and natural enemies in tomato [J]. Ciencia Rural, 2005, 35(1): 204 − 208. doi:  10.1590/S0103-84782005000100033
[72]

SPERANZA S, MELO M C, LUNA M G, et al. First record of Zelus obscuridorsis (Hemiptera: Reduviidae) as a predator of the South American tomato leafminer, Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Florida Entomologist, 2014, 97(1): 295 − 297. doi:  10.1653/024.097.0144
[73] 杨桂群, 范苇, 张倩, 等. 异色瓢虫和龟纹瓢虫幼虫对番茄潜叶蛾低龄幼虫的捕食功能反应[J]. 中国生物防治学报, 2022, 38(4): 959 − 966. doi:  10.16409/j.cnki.2095-039x.2021.09.017
[74]

PICANçO M C, BACCI L, QUEIROZ R B, et al. Social wasp predators of Tuta absoluta [J]. Sociobiology, 2011, 58(3): 621 − 634.
[75]

SIVAKUMAR T, JOSEPHRAJKUMAR A, ANITHA N. First report of tomato pinworm, Tuta absoluta (Meyrick) on egg plant Solanam melongena L. from Kerala, India [J]. Entomon, 2017, 42(4): 335 − 338.
[76]

MOMEN F M, METWALLY A E M, NASR A E K, et al. First report on suitability of the tomato borer Tuta absoluta eggs (Lepidoptera: Gelechiidae) for eight predatory phytoseiid mites (Acari: Phytoseiidae) under laboratory conditions [J]. Acta Phytopathologica et Entomologica Hungarica, 2013, 48(2): 321 − 331. doi:  10.1556/APhyt.48.2013.2.13
[77]

METWALLY A M, MOMEN F M, NASR A K, et al. Prey suitability of Tuta absoluta larvae (Lepidoptera: Gelechiidae) for three predatory phytoseiid mites (Acari: Phytoseiidae) under laboratory conditions [J]. Acta Phytopathologica et Entomologica Hungarica, 2015, 50(1): 105 − 113. doi:  10.1556/038.50.2015.1.10
[78]

ROSTAMI E, MADADI H, ABBASIPOUR H, et al. First report of the predatory spider, Oxyopes lineatus Latreille (Aranea: Oxyopidae) feeding on the tomato leaf miner, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. Advances in Food Sciences, 2018, 40(4): 128 − 133.
[79]

BIONDI A, ZAPPALà L, MAURO A D, et al. Can alternative host plant and prey affect phytophagy and biological control by the zoophytophagous mirid Nesidiocoris tenuis? [J]. BioControl, 2016, 61(1): 79 − 90. doi:  10.1007/s10526-015-9700-5
[80]

MOLLá ó. MONTÓN H, VANACLOCHA P, et al. Predation by the mirids Nesidiocoris tenuis and Macrolophus pygmaeus on the tomato borer Tuta absoluta [J]. IOBC/WPRS Bulletin, 2009, 49: 209 − 214.
[81]

MOLLá ó, GONZALEZ-CABRERA J, URBANEJA A. The combined use of Bacillus thuringiensis and Nesidiocoris tenuis against the tomato borer Tuta absoluta [J]. BioControl, 2011, 56(6): 883 − 891. doi:  10.1007/s10526-011-9353-y
[82]

CALVO F J, LORENTE M J, STANSLY P A, et al. Preplant release of Nesidiocoris tenuis and supplementary tactics for control of Tuta absoluta and Bemisa tabaci in greenhouse tomato [J]. Entomologia Experimentalis et Applicata, 2012, 143: 111 − 119. doi:  10.1111/j.1570-7458.2012.01238.x
[83]

ARNò J, SORRIBAS R, PRAT M, et al. Tuta absoluta, a new pest in IPM tomatoes in the northeast of Spain [J]. IOBC/WPRS Bulletin, 2009, 49: 203 − 208.
[84]

MOAYERI H R S, ASHOURI A, BRøDSGAARD H F, et al. Males of the predatory mirid bug Macrolophus caliginosus exploit plant volatiles induced by conspecifics as a sexual synomone [J]. Entomologia Experimentalis et Applicata, 2007, 123(1): 49 − 55. doi:  10.1111/j.1570-7458.2007.00523.x
[85] 农向群, 张泽华. 昆虫病原真菌的生态适应性及其生物防治应用策略[J]. 中国生物防治学报, 2013, 29(1): 133 − 141. doi:  10.3969/j.issn.2095-039X.2013.01.019
[86]

KLIEBER J, REINEKE A. The entomopathogen Beauveria bassiana has epiphytic and endophytic activity against the tomato leaf miner Tuta absoluta [J]. Journal of Applied Entomology, 2016, 140(8): 580 − 589. doi:  10.1111/jen.12287
[87]

GIUSTOLIN T A, VENDRAMIM J D, ALVES S B, et al. Pathogenicity of Beauveria bassiana (Bals. ) Vuill. to Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) reared on two genotypes of tomato [J]. Neotropical Entomology, 2001, 30(3): 417 − 421. doi:  10.1590/S1519-566X2001000300013
[88]

RODRíGUEZ M, GERDING M, FRANCE A. Entomopathogenic fungi isolates selection for egg control of tomato moth, Tuta absoluta Meyrick (Lepidoptera: Gelechiidae) eggs [J]. Agricultura Tecnica, 2006, 66(2): 151 − 158.
[89]

RODRíGUEZ M, GERDING M, FRANCE A. Effectivity of entomopathogenic fungus strains on tomato moth Tuta absoluta Meyrick (Lepidoptera: Gelechiidae) larvae [J]. Agricultura Tecnica, 2006, 6(2): 159 − 165.
[90]

CONTRERAS J, MENDOZA J E, MARTÍNEZ-AGUIRRE M R, et al. Efficacy of enthomopathogenic fungus Metarhizium anisopliae against Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Journal of Economic Entomology, 2014, 107(1): 121 − 124. doi:  10.1603/EC13404
[91] 关正君, 鲁顺保, 霍艳林, 等. 转Bt基因抗虫作物对非靶标害虫的影响[J]. 生物多样性, 2018, 26(6): 636 − 644. doi:  10.17520/biods.2017315
[92] 张桂芬, 张毅波, 张杰, 等. 苏云金芽胞杆菌G033A对新发南美番茄潜叶蛾的室内毒力及田间防效[J]. 中国生物防治学报, 2020, 36(2): 175 − 183. doi:  10.16409/j.cnki.2095-039x.2020.02.003
[93]

GIUSTOLIN T A, VENDRAMIM J D, ALVES S B, et al. Susceptibility of Tuta absoluta (Meyrick) (Lep., Gelechiidae) reared on two species of Lycopersicon to Bacillus thuringiensis var. kurstaki [J]. Journal of Applied Entomology, 2001, b,125(9/10): 551 − 556.
[94]

GONZALEZ-CABRERA J, MOLLá ó, MONTóN H, et al. Efficacy of Bacillus thuringiensis (Berliner) in controlling the tomato borer, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. BioControl, 2011, 56(1): 71 − 80. doi:  10.1007/s10526-010-9310-1
[95] 刘奇志, 赵映霞, 严毓骅, 等. 我国昆虫病原线虫生物防治应用研究进展[J]. 中国农业大学学报, 2002, 7(5): 65 − 69. doi:  10.3321/j.issn:1007-4333.2002.05.012
[96]

ARTHURS S, HEINZ K M, PRASIFKA J R. An analysis of using entomopathogenic nematodes against above-ground pest [J]. Bulletin of Entomological Research, 2004, 94: 297 − 306. doi:  10.1079/BER2003309
[97]

BATALLA-CARRERA L, MORTON A, GARCíA-DEL-PINO F. Efficacy of entomopathogenic nematodes against the tomato leafminer Tuta absoluta in laboratory and greenhouse conditions [J]. BioControl, 2010, 55(4): 523 − 530. doi:  10.1007/s10526-010-9284-z
[98]

GARCíA-DEL-PINO F, ALABERN X, MORTON A. Efficacy of soil treatments of entomopathogenic nematodes against the larvae, pupae and adults of Tuta absoluta and their interaction with the insecticides used against this insect [J]. BioControl, 2013, 58(6): 723 − 731. doi:  10.1007/s10526-013-9525-z
[99]

DA CUNHA U S, VENDRAMIM J D, ROCHA W C, et al. Potential of Trichilia pallida Swartz (Meliaceae) as a source of substances with insecticidal activity against the tomato leafminer Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. Neotropical Entomology, 2005, 34(4): 667 − 673. doi:  10.1590/S1519-566X2005000400018
[100]

DA CUNHA U S, VENDRAMIM J D, ROCHA W C, et al. Bioactivity of Trichilia pallida Swartz (Meliaceae) derived molecules on Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. Neotropical Entomology, 2008, 37(6): 709 − 715. doi:  10.1590/S1519-566X2008000600013
[101]

MOREIRA M D, PICANçO M C, BARBOSA L C, et al. Toxicity of leaf extracts of Ageratum conyzoides to Lepidoptera pests of horticultural crops [J]. Biological Agriculture & Horticulture, 2004, 22(3): 251 − 260.
[102]

KONA N E M, TAHA A K, MAHMOUD M E E. Effects of botanical extracts of neem (Azadirachta indica) and jatropha (Jatropha curcus) on eggs and larvae of tomato leaf miner, Tuta absoluta (Meyrick) (Lepidoptera: Gelechiidae) [J]. Persian Gulf Crop Protection, 2014, 3(3): 41 − 46.
[103]

DE BRITO E F, BALDIN E L L, SILVA R C M, et al. Bioactivity of Piper extracts on Tuta absoluta (Lepidoptera: Gelechiidae) in tomato [J]. Pesquisa Agropecuaria Brasileira, 2015, 50(3): 196 − 202. doi:  10.1590/S0100-204X2015000300002
[104]

YAROU B B, BAWIN T, BOULLIS A, et al. Oviposition deterrent activity of basil plants and their essentials oils against Tuta absoluta (Lepidoptera: Gelechiidae) [J]. Environmental Science and Pollution Research, 2018, 25(30): 29880 − 29888. doi:  10.1007/s11356-017-9795-6
[105] 陈学新. 21世纪我国害虫生物防治研究的进展、问题与展望[J]. 昆虫知识, 2010, 47(4): 615 − 625.
[106]

CAMARGO R A, BARBOSA G O, POSSIGNOLO I P, et al. RNA interference as a gene silencing tool to control Tuta absoluta in tomato (Solanum lycopersicum) [J]. 2016, PeerJ, 4(12): e2673.
[107]

CAMARGO R A, HERAI R H, SANTOS L N, et al. De novo transcriptome assembly and analysis to identify potential gene targets for RNAi-mediated control of the tomato leafminer (Tuta absoluta) [J]. BMC Genomics, 2015, 16(1): 635. doi:  10.1186/s12864-015-1841-5
[108]

SAVINO V, COVIELLA C E, LUNA M G. Reproductive biology and functional response of Dineulophus phtorimaeae, a natural enemy of the tomato moth, Tuta absoluta [J]. Journal of Insect Science, 2012, 12(153): 1 − 14. doi:  10.1673/031.012.15301
[109] 王伟, 王文霞, 刘万学, 等. 芙新姬小蜂生物学特性及其应用研究进展[J]. 中国生物防治学报, 2012, 28(4): 575 − 582. doi:  10.3969/j.issn.2095-039X.2012.04.020
[110] 薛正帅. 烟盲蝽在生物防治上的研究现状与应用前景[J]. 天津农业科学, 2015, 21(10): 118 − 120. doi:  10.3969/j.issn.1006-6500.2015.10.028
[111] 朱朝东. 中国姬小蜂亚科的系统学研究[D]. 北京: 中国科学院动物研究所, 1998.
[112]

MEDEIROS M A, DE BôAS G L V, VILELA N J, et al. Preliminary survey on the biological control of South American tomato pinworm with the parasitoid Trichogramma pretiosum in greenhouse models [J]. Horticultura Brasileira, 2009, 27(1): 80 − 85. doi:  10.1590/S0102-05362009000100016